Starting with the coherence layer, network operations are simulated in detail, with spiking integrate-and-fire neurons described by
In an integrate-and-fire neuron, the membrane potential develops according to equation (10) as long as , the threshold potential. If is reached, an actionpotential or spike is emitted, and the membrane potential is reset to . After a short refractory period, , where stays at its resting level , the membrane potential develops again according to Equation 10. If a neuron fired at times , it supplies a current of exponentially decaying pulses, , to all postsynaptic neurons.