During dynamical coherence detection, the total output current of the coherence layer is composed of a small random component plus an oscillatory component coming from the synchronous coherence cluster. The oscillatory component has to be detected by the read-out layer .
In the simulations, the read-out neurons
by the total output current of the coherence layer (the first term in equation (13)),
plus a synchronizing interaction term:
The number of read-out neurons ranges between 2 and 16, and the synaptic decay time constant is chosen to equal in the simulations. Network performance turns out to be insensitive to the precise value of , and was set to in all simulations.
The coupling between coherence- and read-out layer, , modifies the capture range of the read-out neurons; in the simulations, it was fixed to , and the synaptic time constant to The results displayed in the maps of the main text were obtained by measuring the spike frequencies of the neurons in the read-out layer and averaging: .